A relatively short (~1.5-3.5 seconds), noisy, pulsating Roaring call by an estrous female when pursued by a male. The Roar of an estrous female often begins as a series of short pulsated growling sounds before developing (or not) into a pulsating Estrous-Roar. This call has the immediate effect of attracting nearby and distant males, or alerting her Consort, or Guarding musth male, who may be dozing or otherwise occupied.
An estrous female is Chased by six males. The largest male is immediately behind her. His running is hampered by his Penis-Erection which is flapping back and forth hitting his legs and we can see him running with his legs wider apart. He attempts to Reach-Over her but is unable to catch her. As he nears her she gives an Estrous-Roar and continues running. He slows and a younger male takes over. (Amboseli, Kenya)
