An estrous female acting vigilant and wary (Chin-Up, Ears-Stiff, Tail-Raising) in the presence of males, gazing at them (Eyes-Wide, Look-At), then orienting away, looking back (Look-Back) over her shoulder to follow their movement, intolerant (Back-Away, Retreat-From or Run-Away, Estrous-Roar) of any male approaches.
Jolie of the JB family is just coming into estrus and she is being followed by Conor in musth. She is exhibiting Wariness - a quickened stride or Purposeful-Walk, Ears-Stiff and Looking-Back to check how close Conor or other males are. (Amboseli, Kenya)