Elephants use Contact Calls to keep in audible contact with
one another sometimes over long distances. In a sense an interchange
of contact calls queries, "I am here, where are you?"
and in answer, "I am over here". Contact calls typically
contain a series of at least three calls: The querying rumble
by the initial caller, an answer by a second individual and
then a confirmation by the initial caller that the answer
has been received(1).
Other nearby family members may also add their voice to the
second or third phase of the series.
Contact calls may be heard relatively frequently in the company
of elephant family groups, particularly when a family has
spread out over a large distance or when it has split into
sub-groups. Contact calls are used primarily between family
and bond group members, but on occasion elephants may be heard
to answer the contact call of a non-family and non-bond-group
elephant(2).
In these cases the calling elephants seem to be members of
the same clan(3).
Contact calls are one of the best studied of the calls made
by savanna elephant(4).
Contact calls can be among the most powerful solo (as opposed
to chorused or clustered calls) low frequency calls made by
elephants. Family members separated by one to two kilometers
use contact calls at high sound pressure levels (measured
at up to 103 dB at 5 m from the source) to stay in contact
with one another(5).
When individuals are closer together, however, they use the
same call type at lower sound pressure levels(6).
It has been estimated that elephants may be able to hear such
powerful calls over an area of 300 km2 under certain atmospheric
conditions(7),
though information regarding individual identity probably
does not travel more than around 2.5 km(8).
Thus, while elephants may be able to identify a call as one
made by a female over distances of up to 10 km, they are probably
only able to use this call for the purposes of staying in
touch with friends and relatives over distances of less than
2.5 km.
Adult females, juveniles and calves all use contact calls.
As far as we are aware adult males do not contact call, but
Joyce's guess is that they probably do when they have the
need to, which is infrequently(9).
Contact calls are typically powerful, loud, throaty, medium-pitched
rumbles that are relatively long in duration. Most contact
calls last between 6-8 seconds and are modulated in frequency
contour, typically rising sharply and then falling gradually.
There does, however, exist a wide range of variation in the
frequency contour of contact calls. Karen McComb and her colleagues(10)
have shown that the contact calls of individuals are structurally
distinct and audibly identifiable to other elephants. In other
words contact calls contain an acoustic signature. It is possible
that considerable additional information is contained in the
different variations of the calls, perhaps related to their
sequential arrangement (i.e. call, answer, confirmation),
or to logistical or locational information or perhaps related
to which individual an elephant is calling. Future research
will answer some of these questions.
Contact calls are quite distinctive both in sound quality
and in behavioral context, but they may on occasion be confused
with the onset of a Greeting Sequence or Greeting Ceremony.
Both the contact rumbles and greeting rumbles are loud modulated
calls and contact calling sometimes precedes a greeting. In
these cases it can be difficult to determine when the contact
calling ends and greeting begins.
The spectrographic examples include a single call and a series.
Note that these are on different time scales. The single call
was one of a long series of calls given by Matriarch Echo
on the 18 August 2000 as she conversed with her age-mate Ella
and Ella's teenage daughter Emma who were some 200 meters
distant.
The other spectrogram depicts a series of calls between a
son, Eldon, and his mother, Eudora. Eldon, a four-year old,
is separated from his mother by about 30 meters and calls
to her rather softly (note that his call overlaps with another
distant but unknown elephant). Eudora answers him and Eldon
responds by calling loudly back to her to which Eudora responds
again.
More information about contact calls see reference (11).